What is the HAO working group?

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The goal of the HAO working group (i.e., the HAO organizing committee and all the morphologists participating in this venture) is to develop an explicit, controlled vocabulary of morphological terms relating to Hymenoptera (the HAO) that:

  • identifies synonyms (homologous structures called different names)
  • textually defines, references, and figures each accepted term with images and annotations
  • establishes and clarifies homologies between hymenopteran lineages
  • is hierarchically arranged (e.g., scape is “part of” the antenna, which is “part of” the head)
  • identifies taxon specific terms (e.g., the cenchrus is found only in “Symphyta”)

Some initial version of the HAO will be approved by the members of the International Society of Hymenopterists (ISH) and made available in a format that allows incorporation into databasing projects. The HAO will also be converted into an online, illustrated atlas of hymenopteran anatomy.


Who is the HAO working group

member email affiliation
Gavin Broad g.broad@nhm.ac.uk Natural History Museum
Andy Deans, chair adeans@gmail.com NC State University
Gary Gibson Gibsong@agr.gc.ca Canadian National Insect Collection
Norm Johnson Johnson.2@osu.edu Ohio State University
Susanne Schulmeister schulmei@amnh.org American Museum of Natural History
Mike Sharkey msharkey@uky.edu University of Kentucky
Bob Wharton rawbaw2@tamu.edu Texas A & M University


Strategy for developing the HAO

The HAO organizing committee will develop the “backbone” of the hierarchy and then solicit volunteers to tackle components of the backbone.

The HymAToL grant is the perfect vehicle for jumpstarting the HAO project in that the hymenopteran body has already been partitioned by lab group for the purpose of coding morphological characters. Each lab group could be responsible for addressing their respective terminology and for drawing in other researchers that are not officially part of HymAToL (to make this a community effort). Current HymAToL assignments are listed below. Each lab would decide on whether to publish their own manuscripts relating to standardizing morphological terms (as well as who should author such publications).


egg unassigned
larva ("Symphyta") unassigned
larva (Apocrita) Dave Wahl (AEI)
adult head Heraty lab (UC-Riverside)
adult mesosoma (thorax) Ronquist lab (FSU)
adult legs Ronquist lab (FSU)
wings Sharkey lab (UKY)
metasoma (abdomen) unassigned
female genitalia Sharkey lab (UKY)
male genitalia Schulmeister (AMNH)
surface sculpturing Deans et al. (FSU)
propodeal carinae Dave Wahl (AEI)
other terms* unassigned

*e.g., meconium, setae, sensilla, pores/glands, line vs. sulcus vs. carina, etc.

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